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Contact Institute of Immunology and Infection Research University of Edinburgh Ashworth Labs King's Buildings West Mains Road Edinburgh EH9 3JT Scotland Tel: 0131 6507573 Fax: 0131 6506564 nick.savill@ed.ac.uk		Affiliations Centre for Infection, Immunity and Evolution (CIIE) Edinburgh Infectious Diseases School of Biological Sciences University of Edinburgh IIIR